李丽霞,冯洪真,李明,彭进,季鑫鑫,王文卉

• 1:南京大学地球科学系
• 2:中国地质科学院地质研究所

摘要(Abstract)：

根据对采自湖南益阳南坝剖面的笔石化石的研究,识别出了下奥陶统弗洛阶4个连续的笔石带(自下而上):Tetragraptus approximatus带、Pendeograptus fruticosus带、Didymograptellus bifidus带和Corymbograptus deflexus带。同时在特马豆克阶顶部也识别出了Hunnegraptus copiosus带,从而在该区确立了一个连续的、可与国内外同期地层精确对比的弗洛阶笔石带序列。

关键词(KeyWords)： 笔石;生物带;弗洛阶;下奥陶统;益阳;湖南

Abstract:

Keywords:

基金项目(Foundation): 国家自然科学基金(No.40472014)资助项目

作者(Author): 李丽霞,冯洪真,李明,彭进,季鑫鑫,王文卉

参考文献(References)：

• 陈旭,韩乃仁.1964.江西玉山早奥陶世笔石地层.地质论评,22(2):81—90
• 陈旭,戎嘉余,汪啸风,周志毅,王志浩,陈挺恩,耿良玉,邓占球,胡兆,董得源,李军,张元动,詹仁斌.1993.中国奥陶纪生物地层学研究的新进展.地层学杂志,17(2):89—99
• 陈旭,杨达铨.1988.江西玉山早奥陶世笔石动物群的分带及分异.地层学杂志,12(2):112—124
• 陈旭,杨达铨,韩乃仁,李罗照.1983.江西玉山下奥陶统宁国组底部工字笔石带的笔石.古生物学报,22(3):324—330
• 陈旭,杨万蓉,何自强,汪盛辉.1981.广西兴安奥陶纪含笔石地层.地层学杂志,5(1):34—43
• 陈旭,张元动,樊隽轩.2006.奥陶纪笔石的主要演化辐射进程.见:生物的起源、辐射与多样性演变——华夏化石记录的启示.北京:科学出版社.181—196
• 陈旭,周志毅,戎嘉余,李军.2000.奥陶系.见:中国科学院南京地质古生物研究所编.中国地层研究二十年(1979—1999).合肥:中国科学技术大学出版社.39—58
• 陈旭,Bergstr m S M.2008.奥陶系研究百余年:从美国标准到国际标准.地层学杂志,32(1):1—14
• 李积金,陈旭.1962.黔南三都寒武纪及奥陶纪笔石.古生物学报,10(1):12—35
• 黎作聪.1980.鄂西下奥陶统大湾组下部的下垂和下曲类对笔石动物群.湖北区测——地层古生物专集.46—56
• 穆恩之.1959.中国含笔石地层——中国地质学基本资料专题总结论文集第3号.北京:地质出版社.1—75
• 穆恩之,葛梅钰,陈旭,倪寓南,林尧坤.1979.西南地区下奥陶统的笔石.中国古生物志,总号第156册,新乙种第13号.北京:科学出版社.1—192
• 穆恩之,李积金,葛梅钰,林尧坤,倪寓南.2002.中国笔石.北京:科学出版社.1—1042
• 汪啸风,金玉琴,吴兆同,傅汉英,黎作聪,马国干.1978.笔石纲.见:湖北省地质科学研究所等.中南地区古生物图册(一)早古生代部分.北京:地质出版社.266—371
• 汪啸风,Stouge S.,陈孝红,李志宏,王传尚,Erdt mann B-D,曾庆銮,周志强,陈辉明,张淼,徐光洪.2005.全球下奥陶统-中奥陶统界线层型候选剖面——宜昌黄花场剖面研究新进展.地层学杂志,29(3):467—489
• 肖承协,陈洪冶.1990.玉山古城一带早中奥陶世笔石动物群.江西地质,4(2):83—243
• 肖承协,薛春汀,黄学涔.1975.江西崇义早奥陶世笔石地层.地质学报,49(2):112—125
• 许杰.1934.长江下游之笔石化石.见:许杰笔石论文集.北京:地质出版社.1—108
• 许杰,黄枝高.1979.新疆霍城县果子沟地区下奥陶统的笔石动物群.地质学报,53(1):1—19
• 张元动,陈旭.2006.华南早—中奥陶世笔石动物群的辐射演化.见:戎嘉余主编.生物的起源、辐射与多样性演变——华夏化石记录的启示.北京:科学出版社.285—316
• 张元动,刘晓,詹仁斌.2007.贵州遵义高桥奥陶系湄潭组下部的笔石.古生物学报,46(2):145—166
• 张元动,王志浩,冯洪真,骆天天,Erdt mann B-D.2005.中国特马豆克阶笔石地层评述.地层学杂志,29(3):215—235
• Albanesi G L,Ortega G,Barnes C R&Hunicken M A.1999.Co-nodont-graptolite biostratigraphy of the Gualcamayo Formation(Middle Ordovician)in the Gualcamayo-Guandacol rivers area,Argentina Precordillera.Acta Universitatis Carolinae-Geologi-ca,43(1-2):45—48
• Albanesi G L,Carrera M G,Ca灳as F L&Saltzman M.2003.TheNiquivil section,Precordillera of San Juan,Argentina,pro-posed GSSP for Lower/Middle Ordovician boundary.In:Al-banesi G L,Beresi M S&Peralta S H eds.Ordovician fromAndes.INSUGEO(Serie Correlaci幃n Geol幃gica),17:33—40
• Bergstr m S M&Chen Xu.2007.Ordovician Correlation Chartcompiled by Stig Bergstr mand Chen Xu.In:ISOS,Website
• Bergstr mS M,Finney S C,Chen Xu,Goldman D&Leslie S A.2006.Proposed names for three Ordovician global stages.Or-dovician News,23:10—11
• ergstr mS M,L fgren A&Maletz J.2004.The GSSP of theSecond(Upper)Stage of the Lower Ordovician Series:Diabas-brottet at Hunneberg,Province of Vasterg tland,Southwest-ern Sweden.Episodes,27(4):265—272
• Berry WB N.1962.Graptolite occurrence and ecology.Journal ofPaleontology,36(2):285—293
• Chen Xu&Han Nai-ren.1964.The early Ordovician biostratigra-phy of graptolites in Yushan,Jiangxi.Geological Review,22(2):81—90
• Chen Xu,Rong Jia-yu,Wang Xiao-feng,Zhou Zhi-yi,Wang Zhi-hao,Chen Ting-en,Geng Liang-yu,Deng Zhan-qiu,Hu Zhao-xun,Dong De-yuan,Li Jun,Zhang Yuan-dong&Zhan Ren-bin.1993.Newadvances in the study of Ordovician biostratig-raphy in China.Journal of Stratigraphy,17(2):89—99
• Chen Xu,Rong Jia-yu&Zhou Zhi-yi.2003.Ordovician biostratig-raphy of China.In:Zhang Wen-tang,Chen Pei-ji&Pal mer AReds.Biostratigraphy of China.Beijing:Science Press.121—171
• Chen Xu&Yang Da-quan.1988.Biozonation and diversification ofearly Ordovician graptolite fauna from Yushan,Jiangxi.Jour-nal of Stratigraphy,12(2):112—124
• Chen Xu,Yang Da-quan,Han Nai-ren&Li Luo-zhao.1983.Grap-tolites fromthe Tetragraptus(Etagraptus)approxi matus zoneof the lowermost Ningguo Formation in Yushan,NE Jiangxi.Acta Palaeontologica Sinica,22(3):324—330
• Chen Xu,Yang Wan-rong,He Zi-qiang&Wang Sheng-hui.1981.Ordovician graptolite bearing beds from Xingan,Guangxi.Journal of Stratigraphy,5(1):34—43
• Chen Xu,Zhang Yuan-dong&Fan Jun-xuan.2006.A brief intro-duction to the evolutionary radiation of Ordovician graptolites.In:Originations,radiations and biodiversity changes-evidencesfromthe Chinese fossil record.Beijing:Science Press.181—196
• Chen Xu,Zhou Zhi-yi,Rong Jia-yu&Li Jun.2000.Ordovician.In:NanjingInstitute of Geology and Palaeontology,Chinese A-cademy of Sciences ed.Stratigraphy studies of China for20years(1979—1999).Hefei:University of Science and Technol-ogy of China Press.39—58
• Chen Xu&Wang Zhi-hao.1993.The base of Arenig series in Chi-na.Newsletters on Stratigraphy,29(3):159—164
• Cooper R A&Fortey R A.1982.The Ordovician graptolites oSpitsbergen.Bulletin of the British Museum Natural Histor(Geology Series),36:157—302
• Cooper R A&Lindhol m M.1990.Aprecise worldwide correlationof early Ordovician graptolite sequence.Geological Magazine,127(6):497—525
• Cooper A H,Rushton A WA,Molyneux S G,Hughes R A,MooreR M&Webb B C.1995.The stratigraphy,correlation,prove-nance and palaeogeogeaphy of the Skiddaw Group(Ordovician)in the English Lake District.Geological Magazine,132(2)185—211
• Cooper R A,Nowlan GS&Williams S H.2001.Global StratotypeSection and Point for base of the Ordovician System.Episodes,24(1):19—28
• Egenhoff S O,Maletz J&Erdt mann B-D.2004.Lower Ordoviciangraptolite biozone and lithofacies of southern Bolivia:relevancefor palaeogeographicinterpretations.Geological Magazine,141(3):287—299
• Elles G L&Wood E M R.1901.A monograph of British grapto-lites,Part1-Dichograptidae.Palaeontographical SocietyLon-don,1—54
• Feng Hong-zhen,Erdt mann B-D,Li Ming,Peng Jin,Li Li-xia&JiXin-xin.2007.Newdata on upper Tremdocian to lower Floiangraptolite succession in south China.Acta Palaeontologica Si-nica,46(Suppl.):120—127
• Finney S C.1982.Ordovician graptolite zonation.In:Ross RJ Jretal.The Ordovician Systemin the United States.Correlationchart and explanatory notes.International Union of GeologicalScience,Publication12:14—23
• Finney S C&Ethington R L.1992.Whiterockian graptolites andconodonts fromthe Vinini Formation,Nevada:biostratigraphici mplications.In:Webby B D&Laurie J Reds.Global perspec-tives on Ordovician geology.Rotterdam:Balkema A A.153—169
• Hall J.1858.Descriptions of Canadian graptolites.Geological Survey of Canada,Report for1857:111—145
• Jackson D E&Lenz A C.2003.Taxonmic and biostratigraphic sig-nificance of the Tremadoc graptolite fauna fromnorthern YukonTerritory,Canada.Geological Magazine,104(2):131—156
• Li Ji-jin&Chen Xu.1962.Cambrian and Ordovician graptolitesfromSandu,S.Guizhou(Kuichou).Acta Palaeontologica Sin-ica,10(1):12—35
• Li Zuo-cong.1980.The pendent and deflexed didymograptid faunafromthe lower part of Dawan Formation(Lower Ordovician),western Hubei.Geologcial Survey of Hubei,Stratigraphyand Palaeontology,Special Volume46—56
• Mu En-zhi.1959.Graptolite bearing strata of China:Chinese geolo-gy basicinformation special summarize collection of theses(No.3).Beijing:Geological Publishing House.1—75
• Mu En-zhi,Ge Mei-yu,Chen Xu,Ni Yu-nan&Lin Yao-kun.1979.Lower Ordovician graptolites of Southwest China.Palae-ontological Sinica,Whole No.156(New Series B,No.13).Beijing:Science Press.1—192
• Mu En-zhi,Li Ji-jin,Ge Mei-yu,Lin Yao-kun&Ni Yu-nan.2002.Fossil graptolites of China.Beijing:Science Press.1—1042
• Maletz J.1994.Pendent didymograptids(Graptoloidea,Dichograp-tacea).In:Chen Xu,Erdt mann B-D&Ni Yu-nan eds.Grapto-lite research today.Nanjing:Nanjing University Press.27—43
• Maletz J,L fgren A&Bergstr m S M.1996.Proposal for the a-doption of the Diabasbrottet section(Hunneberg,Vasterg tland)as a Global stratotype section and point(GSSP)for the second series of the Ordovician System.Newsletters onStratigraphy,34:129—159
• Maletz J.1997.Arenig biostratigraphy of the Point-de-Lvy slice,Quebec Appalachians,Canada.Canadian Journal of EarthScience,34:733—752
• Maletz J&Egenhoff S O.2001.Late Tremadoc to early Areniggraptolite faunas of southern Bolivia and their i mplications for aworldwide biozonation.Lethaia,34:47—62
• Mosen A.1937.Die graptolithenfauna i munterenDidymograptus-schiefer(Phyllograptusschiefer)Norwegens.Norsk Geolo-gisk Tiddsskrift,16:57—266
• Mu En-zhi.1974.Evolution,classification and distribution of Grap-toidea and graptodendroids.Scientia Sinica,17:227—238
• Nicholson H A.1873.On some fossils from the Quebec group ofPoint Lv s,Quebec.Annals and Magazine of Natural Histo-ry,4(11):133—143
• Ruedemann R.1947.Graptolites of North America.Geological So-ciety of America,Memoir19:1—652
• Tol macheva T&Fedorov P.2001.The Ordovician Billingen/Volkhov boundaryinterval(Arenig)at Lava River,northwest-ern Russia.Norwegian Journal of Geology,81:161—168
• Tzaj.1974.Lower Odovician graptolites of Kazakhstan.Moscow:Nauka.1—127(in Russian)
• Tullberg S A.1880.NagraDidymograptus-artus i undre grap-tolitskiffer vid Kiviks-Esper d.Geologiska F reningens i Stock-hol m F rhandlingar,5(58)(2):39—43(in Swedish)
• VandenBerg A H M&Cooper R A.1992.The Ordovician grapto-lite sequence of Australasia.Alcherina,16:33—85
• Wang Xiao-feng,Jin Yu-qin,Wu Zhao-tong,Fu Han-ying,Li Zuo-cong&Ma Guo-gan.1978.Graptolithina.In:Hubei Instituteof Geoscienceset.al.eds.A palaeontological atlas of centralsouth China(Early Palaeozoic).Beijing:Geological PublishingHouse.266—371
• Wang Xiao-feng,Li Zhi-hong,Chen Xiao-hong&Wang Chuan-shang.2003.The Huanghuachang section,potential as GlobalStratotype for the base of the Middle Ordovician Series.In:Al-banesi G L,Beresi M S&Peralta S H eds.Ordovician fromAndes.INSUGEO,Serie Correlacin Geolgica17:153—159
• Wang Xiao-feng,Stouge S,Chen Xiao-hong,Li Zhi-hong,WangChuan-shang,Erdt mann B-D,Zeng Qing-luan,Zhou Zhi-qiang,Chen Hui-ming,Zhang Miao&Xu Guang-hong.2005.Advances on the potential GSSPfor the base of Middle Ordovi-cian series-Huanghuachang Section.Journal of Stratigraphy,29(3):467—489
• Wang Xiao-feng,Stouge S,Erdt mann B-D,Chen Xiao-hong,LiZhi-hong,Wang Chuan-shang,Zeng Qing-luan,Zhou Zhi-qiang&Chen Hui-ming.2005.Aproposed GSSPfor the base of theMiddle Ordovician Series:the Huanghuachang section,Yichang,China.Episodes,28(2):105—117
• Wang Xiao-feng&Wang Chuan-shang.2001.Tremadocian(Ordo-vician)graptolite diversification events in China.Alcheringa,25:155—168
• Webby B D.1996.ISOS annual report for1995.Ordovician News,13:2—7
• Webby B D,Cooper R A,Bergstr mS M&Paris F.2004.Strati-graphic framework and ti me slices.In:Webby B D,Paris F,Droser ML&Percival I eds.The great Ordovician biodiversifi-cation event.New York:Columbia University Press.41—47
• Williams S H&Stevens R K.1988.Early Ordovician(Arenig)graptolites of the Cow Head Group,western Newfoundland,Canada.Palaeontologica Canadiana,5:1—167
• Williams S H,Barnes C R,O'Brien F HC&Boyce WD.1994.Aproposed global stratotype for the Second Series of the Ordovi-cian System:Cow Head Peninsula,western Newfoundland.Bulletin of Canadian Petroleum Geology,42:219—231
• Williams S H&Barnes C R.1996.Discussion of the Ledge section,western Newfoundland,as a suitable GSSP section for an earlyOrdovician Stage/Subseries boundary.Ordovician News,13:47—53
• Xiao Cheng-xie&Chen Hong-ye.1990.Early and Middle Ordovi-cian graptolite fauna from Gucheng of Yushan.Geology ofJiangxi,4(2):83—243
• Xiao Cheng-xie,Xue Chun-ting&Huang Xue-cen.1975.Early Or-dovician graptolite bearing beds from Chongyi,Jiangxi.ActaGeologica Sinica,49(2):112—125
• Xu Jie.1934.The graptolites of the Lower Yangtze Valley.In:Graptolites collected papers of Xu Jie.Beijing:Geological Pub-lishing House.1—108
• Xu Jie&Huang Zhi-gao.1979.Lower Ordovician graptolite faunaof Guozigou area,Huocheng,Xinjiang.Acta Geologica Sinica,53(1):1—19
• Zhang Yuan-dong&Chen Xu.2003.The Early-Middle Ordoviciangraptolite sequence of the Upper Yangtze region,South China.In:Albanesi G L,Beresi MS&Peralta S Heds.Ordovicianfrom Andes.INSUGEO,Serie Correlacin Geolgica17:173—180
• Zhang Yuan-dong&Chen Xu.2006.Evolutionary radiation of theEarly-Mid Ordovician graptolites in South China.In:Origina-tions,radiations and biodiversity changes:evidences from theChinese fossil record.Beijing:Science Press.285—316
• Zhang Yuan-dong,Erdt mann B-D&Feng Hong-zhen.2004.Tre-madocian(Early Ordovician)graptolite biostratigraphy of Chi-na.Newsletter on Stratigraphy,40:155—182
• Zhang Yuan-dong,Liu Xiao&Zhan Ren-bin.Early and Middle Or-dovician graptolites from the Meitan Formation in Zunyi,Guizhou,China.Acta Palaeontologica Sinica,46(2):145—166
• Zhang Yuan-dong,Wang Zhi-hao,Feng Hong-zhen,Luo Tian-tian&Erdt mann B-D.2005.Tremadocian(Ordovician)graptolitebiostratigraphy of China:Areview.Journal of Stratigraphy,29(3):215—235